As you said it works with default template, you need to check what has changed in your custom template
-- The item list is created by category_items.php template file and not be category.php template file
-- inside the category_items.php template file the lines that link title use parameter 'link_titles' search for it
-- since your template is a copy of default check what you have changed in it, maybe manually recopy the category_items.php from default template to your custom template,
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